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Caractères dérivés propres :
– La carapace dorsale est spécifiquement constitué d’os plats soudés les uns aux autres, surmontés d’écailles épaisses. Les os plats sont liés aux côtes et aux vertèbres.
– Le ventre est protégé par une carapace ou plastron
– Un bec corné
– Les maxillaires, prémaxillaires et dentaire sont dépourvus des dents
– Perte de la double fenestration temporale
– Perte de l’os post-temporal
– Perte du cleithrum (os de la ceinture scapulaire)
– Dix vertèbres du tronc
– Les ceintures sont à l’intérieur de la cage thoracique